#METABOLOMICS WORKBENCH Mahanlab_2025_20250703_231533 DATATRACK_ID:6133 STUDY_ID:ST004042 ANALYSIS_ID:AN006681 PROJECT_ID:PR002534
VERSION                          	1
CREATED_ON                       	08-04-2025
#PROJECT
PR:PROJECT_TITLE                 	Untargeted metabolomics analysis on mouse hepatocytes
PR:PROJECT_SUMMARY               	To determine whether specific metabolites are associated with the induction of
PR:PROJECT_SUMMARY               	P2-HNF4α in liver disease and hepatocellular carcinoma (HCC), we performed
PR:PROJECT_SUMMARY               	untargeted metabolomics on hepatocytes isolated from wild-type mouse liver,
PR:PROJECT_SUMMARY               	which predominantly express the tumor-suppressive P1-HNF4α isoform. Using
PR:PROJECT_SUMMARY               	lentiviral transduction, we overexpressed the P2-HNF4α isoform and conducted
PR:PROJECT_SUMMARY               	metabolite profiling on cell lysates from both control and
PR:PROJECT_SUMMARY               	P2-HNF4α-overexpressing cells. Data analysis was performed using MetaboAnalyst
PR:PROJECT_SUMMARY               	(v5.0), resulting in a list of approximately 120 matched compounds representing
PR:PROJECT_SUMMARY               	significantly predicted metabolites. This study aims to uncover metabolic
PR:PROJECT_SUMMARY               	signatures associated with P2-HNF4α expression, offering insights into its
PR:PROJECT_SUMMARY               	potential role in liver pathogenesis and tumor progression.
PR:INSTITUTE                     	University of Texas Health Science Center at Houston
PR:DEPARTMENT                    	IMM
PR:LABORATORY                    	Mahan Lab
PR:LAST_NAME                     	Fekry
PR:FIRST_NAME                    	Baharan
PR:ADDRESS                       	1825 Pressler St, Houston TX 77030. USA,
PR:EMAIL                         	Baharan.Fekry@uth.tmc.edu
PR:PHONE                         	7135002487
PR:DOI                           	http://dx.doi.org/10.21228/M83G11
#STUDY
ST:STUDY_TITLE                   	Untargeted Metabolomics Identifies Oncometabolites Induced by P2-HNF4α
ST:STUDY_TITLE                   	Overexpression in Hepatocytes
ST:STUDY_SUMMARY                 	This study explores the metabolic impact of P2-HNF4α overexpression in primary
ST:STUDY_SUMMARY                 	mouse hepatocytes. Using untargeted metabolomics, we compared metabolite
ST:STUDY_SUMMARY                 	profiles between control and P2-HNF4α-overexpressing cells to identify
ST:STUDY_SUMMARY                 	metabolic pathways potentially altered in liver disease and HCC.
ST:INSTITUTE                     	University of Tennessee Health Science Center
ST:DEPARTMENT                    	IMM
ST:LABORATORY                    	Mahan Lab
ST:LAST_NAME                     	Fekry
ST:FIRST_NAME                    	Baharan
ST:ADDRESS                       	1825 Pressler St, Houston TX 77030. USA.
ST:EMAIL                         	Baharan.Fekry@uth.tmc.edu
ST:PHONE                         	18434693199
ST:SUBMIT_DATE                   	2025-07-03
#SUBJECT
SU:SUBJECT_TYPE                  	Cultured cells
SU:SUBJECT_SPECIES               	Mus musculus
SU:TAXONOMY_ID                   	10090
#SUBJECT_SAMPLE_FACTORS:         	SUBJECT(optional)[tab]SAMPLE[tab]FACTORS(NAME:VALUE pairs separated by |)[tab]Additional sample data
SUBJECT_SAMPLE_FACTORS           	-	HEP_6FP1	Sample source:wildtype | Treatment:GFP	
SUBJECT_SAMPLE_FACTORS           	-	HEP_6FP2	Sample source:wildtype | Treatment:GFP	
SUBJECT_SAMPLE_FACTORS           	-	HEP_6FP3	Sample source:wildtype | Treatment:GFP	
SUBJECT_SAMPLE_FACTORS           	-	HEP_A8_1	Sample source:wildtype | Treatment:P2_HNF4a	
SUBJECT_SAMPLE_FACTORS           	-	HEP_A8_2	Sample source:wildtype | Treatment:P2_HNF4a	
SUBJECT_SAMPLE_FACTORS           	-	HEP_A8_3	Sample source:wildtype | Treatment:P2_HNF4a	
#COLLECTION
CO:COLLECTION_SUMMARY            	Hepatocytes were isolated from mouse livers and transduced in culture with
CO:COLLECTION_SUMMARY            	either GFP or P2-HNF4α. Following treatment, cells were collected and subjected
CO:COLLECTION_SUMMARY            	to untargeted metabolomics analysis.
CO:SAMPLE_TYPE                   	Cultured cells
#TREATMENT
TR:TREATMENT_SUMMARY             	Primary hepatocytes were isolated from mouse livers and cultured in vitro. Cells
TR:TREATMENT_SUMMARY             	were transduced with either control GFP or P2-HNF4α-expressing lentivirus.
TR:TREATMENT_SUMMARY             	Following 48 hours of transduction, cells were harvested and processed for
TR:TREATMENT_SUMMARY             	untargeted metabolomics analysis.
#SAMPLEPREP
SP:SAMPLEPREP_SUMMARY            	Samples were diluted in a solution containing 25% acetonitrile, 75% water, and
SP:SAMPLEPREP_SUMMARY            	0.1% formic acid prior to LC-MS analysis. The diluted samples were injected into
SP:SAMPLEPREP_SUMMARY            	an Agilent 1200 Series HPLC system coupled to an Agilent 6538 UHD Accurate-Mass
SP:SAMPLEPREP_SUMMARY            	Q-TOF mass spectrometer. Chromatographic separation was performed using an
SP:SAMPLEPREP_SUMMARY            	Agilent ZORBAX Extend-C18 Rapid Resolution HT column (2.1 × 50 mm, 1.8 µm)
SP:SAMPLEPREP_SUMMARY            	under a 20-minute gradient of 25% to 90% Buffer B (80% acetonitrile, 20% water,
SP:SAMPLEPREP_SUMMARY            	0.1% formic acid) at a flow rate of 0.2 mL/min.
#CHROMATOGRAPHY
CH:CHROMATOGRAPHY_SUMMARY        	Liquid chromatography was performed using an Agilent 1200 Series HPLC system
CH:CHROMATOGRAPHY_SUMMARY        	equipped with an Agilent ZORBAX Extend-C18 Rapid Resolution HT column (2.1 × 50
CH:CHROMATOGRAPHY_SUMMARY        	mm, 1.8 µm). The mobile phases consisted of Buffer A (2% acetonitrile, 98%
CH:CHROMATOGRAPHY_SUMMARY        	water, 0.1% formic acid) and Buffer B (80% acetonitrile, 20% water, 0.1% formic
CH:CHROMATOGRAPHY_SUMMARY        	acid). A linear gradient from 25% to 90% Buffer B was applied over 20 minutes at
CH:CHROMATOGRAPHY_SUMMARY        	a flow rate of 0.2 mL/min to achieve metabolite separation prior to MS analysis.
CH:INSTRUMENT_NAME               	Agilent 1200 Series HPLC coupled to Agilent 6538 UHD Accurate-Mass Q-TOF
CH:COLUMN_NAME                   	Agilent ZORBAX Extend-C18 Rapid Resolution HT (2.1 × 50 mm, 1.8 µm)
CH:COLUMN_TEMPERATURE            	21°C
CH:FLOW_GRADIENT                 	25% to 90% Solvent B over 20 minutes
CH:FLOW_RATE                     	0.2 mL/min
CH:SOLVENT_A                     	2% Acetonitrile/98% Water; 0.1% formic acid
CH:SOLVENT_B                     	80% Acetonitrile/20% Water; 0.1% formic acid
CH:CHROMATOGRAPHY_TYPE           	Reversed phase
#ANALYSIS
AN:ANALYSIS_TYPE                 	MS
#MS
MS:INSTRUMENT_NAME               	Agilent 6538 QTOF
MS:INSTRUMENT_TYPE               	QTOF
MS:MS_TYPE                       	ESI
MS:MS_COMMENTS                   	MS1 spectra were acquired over a mass range of 100–2000 m/z using electrospray
MS:MS_COMMENTS                   	ionization (ESI) in positive mode. Acquisition was performed under the following
MS:MS_COMMENTS                   	conditions: capillary voltage of 3.5 kV, drying gas flow rate of 5 L/min,
MS:MS_COMMENTS                   	nebulizer pressure of 5 psi, and source temperature of 300°C.
MS:ION_MODE                      	POSITIVE
#MS_METABOLITE_DATA
MS_METABOLITE_DATA:UNITS         	LC/MS Peak intensity
MS_METABOLITE_DATA_START
Samples	HEP_6FP1	HEP_6FP2	HEP_6FP3	HEP_A8_1	HEP_A8_2	HEP_A8_3
Factors	Sample source:wildtype | Treatment:GFP	Sample source:wildtype | Treatment:GFP	Sample source:wildtype | Treatment:GFP	Sample source:wildtype | Treatment:P2_HNF4a	Sample source:wildtype | Treatment:P2_HNF4a	Sample source:wildtype | Treatment:P2_HNF4a	
11_12_15-THETA			0.0007	0.0004		0.0003
11_14_15-THETA			0.0008	0.0004		0.0003
11b_17a_21-Trihydroxypreg-nenolone			0.0011			
11b_21-Dihydroxy-3_20-oxo-5b-pregnan-18-al			0.0013			
11-cis-Retinaldehyde		0.0003				
11-cis-Retinol	0.0010		0.0003			
11H-14_15-EETA			0.0014			
1_2-Dihydronaphthalene-1_2-diol		0.0004				
12(R)-HPETE			0.0006			
12(S)-HPETE			0.0003			
15H-11_12-EETA			0.0007			
15(S)-HPETE			0.0000			
16a-Hydroxyandrost-4-ene-3_17-dione			0.0009		0.0010	
16b-Hydroxyestrone			0.0004			
17alpha_21-Dihydroxypregnenolone			0.0002			
17-beta-Estradiol-3-glucuronide			0.0007		0.0005	0.0004
17-Hydroxyprogesterone					0.0010	
19-Hydroxyandrost-4-ene-3_17-dione			0.0004		0.0010	
19-Oxoandrost-4-ene-3_17-dione			0.0006		0.0005	
19-Oxotestosterone			0.0006		0.0010	
1a_11b-Dihydro-4_9-dimethylbenz[a]anthra[3_4-b]oxirene			0.0014			
(1aalpha_2beta_3alpha_11calpha)-1a_2_3_11c-Tetrahydro-6_11-dimethylbenzo[6_7]phenanthro[3_4-b]oxirene-2_3-diol				0.0013	0.0002	
1-Methylxanthine	0.0002					
1-Naphthol		0.0004			0.0006	
1-Nitronaphthalene-5_6-oxide					0.0002	
1-Nitronaphthalene-7_8-oxide					0.0005	
1-Pyrroline-4-hydroxy-2-carboxylate				0.0001		0.0004
20a-Dihydroprogesterone		0.0007				
20-Hydroxy-leukotriene B4			0.0010	0.0012		
21-Deoxycortisol			0.0017			
24-Hydroxycalcitriol		0.0004				
2-Hydroxy-3-(4-hydroxyphenyl)propenoic acid		0.0002	0.0000			
2-Hydroxyestrone			0.0004			
2-Methoxyestradiol			0.0008		0.0010	
2-Methoxy-estradiol-17b 3-glucuronide				0.0011		
2-Methoxyestrone			0.0006		0.0010	
2-Oxo-4-methylthiobutanoic acid			0.0001		0.0003	
2-Oxoarginine		0.0000	0.0000			
2-(S-Glutathionyl)acetyl glutathione					0.0002	
3_4-Dihydroxyphenylacetaldehyde			0.0003			
3_4-Dihydroxyphenylglycol	0.0007				0.0004	
3a_21-Dihydroxy-5b-pregnane-11_20-dione			0.0015			
3a_7a_12a-Trihydroxy-5b-cholestan-26-al				0.0009		
3a_7a-Dihydroxycoprostanic acid				0.0005		
3beta_7alpha-Dihydroxy-5-cholestenoate		0.0004				
3beta-Hydroxypregn-5-en-20-one sulfate				0.0002	0.0005	
3-Hydroxylidocaine			0.0006		0.0001	
3-ketosphinganine					0.0185	0.0001
3-Methoxy-4-hydroxyphenylglycolaldehyde		0.0003	0.0001			
4a-Carboxy-4b-methyl-5a-cholesta-8_24-dien-3b-ol				0.0002		
4a-Hydroxytetrahydrobiopterin				0.0002		
4-Guanidinobutanoic acid			0.0004			
4-Hydroxy-4-(methylnitrosoamino)-1-(3-pyridinyl)-1-butanone						0.0005
4-Hydroxydebrisoquine			0.0001		0.0008	
4-[(Hydroxymethyl)nitrosoamino]-1-(3-pyridinyl)-1-butanone						0.0003
4-Hydroxyphenylacetaldehyde		0.0002	0.0003			
4-Hydroxyphenylpyruvic acid		0.0002	0.0000			
4-Hydroxy tolbutamide	0.0012		0.0004	0.0012	0.0001	0.0012
4-Pyridoxic acid						0.0012
4-Trimethylammoniobutanoic acid		0.0006				
5_10-Methylene-THF			0.0010			
5_6-Dihydro-5-fluorouracil					0.0003	
5_7_24(28)-Ergostatrienol			0.0003			
5-Acetylamino-6-formylamino-3-methyluracil		0.0000				
5a-Dihydrotestosterone sulfate	0.0005	0.0003				0.0014
5-alpha-Dihydrotestosterone glucuronide	0.0020	0.0000	0.0021			0.0016
5-Androstenediol			0.0009			
5a-Pregnane-3_20-dione		0.0003				
5b-Dihydrotestosterone			0.0011			
5-Fluorouridine				0.0004	0.0007	
5-Formiminotetrahydrofolic acid				0.0003		
5-Hydroxykynurenine					0.0004	
5-L-Glutamyl-taurine			0.0003			
5-Methoxyindoleacetate			0.0004			
5-Methyltetrahydrofolic acid	0.0023		0.0013			
5^-Methylthioadenosine					0.0001	
5-phosphonooxy-L-lysine					0.0006	
5(S)-Hydroperoxyeicosatetraenoic acid			0.0000			
6-Hydroxymelatonin						0.0001
6-Methylthiopurine			0.0004			
7_12-Dimethylbenz[a]anthracene 5_6-oxide			0.0013			
7_8-Dihydroneopterin			0.0003			
7alpha-Hydroxy-3-oxo-4-cholestenoate			0.0013			
7-Hydroxymethyl-12-methylbenz[a]anthracene			0.0014			
7-Hydroxymethyl-12-methylbenz[a]anthracene sulfate					0.0008	
7-Methylxanthine	0.0002					
8-Isoprostaglandin F2a	0.0015		0.0004	0.0006		0.0003
8(S)-HPETE			0.0003			
9-cis-Retinal		0.0003				
9-cis-Retinoic acid					0.0002	
9-cis-Retinol	0.0010		0.0003			
9-Hydroxybenzo[a]pyrene						0.0006
Acetylcholine chloride		0.0007				
Acetyl-CoA				0.0005		
Adenosine		0.0013	0.0004		0.0012	
ADP-ribose 2^-phosphate			0.0024			
Adrenic acid				0.0007		
Adrenosterone			0.0004		0.0009	
Aflatoxin B1					0.0005	
all-trans-Decaprenyl diphosphate			0.0004			
All-trans-retinoic acid					0.0001	
alpha-Fluoro-beta-ureidopropionic acid					0.0003	
Alpha-Lactose					0.0013	
Alpha-Tocotrienol		0.0007	0.0014			0.0012
Aminoadipic acid		0.0000				
Androstanedione					0.0002	
Androstenedione			0.0008			
Androsterone			0.0010			
Androsterone glucuronide			0.0002			
Anserine		0.0000			0.0002	
Arachidic acid				0.0004		
Arachidonic acid			0.0015			
Argininosuccinic acid	0.0006					
B-Carotene			0.0018			
Benzo[a]pyrene						0.0005
Benzo[a]pyrene-4_5-oxide						0.0006
Benzo[a]pyrene-7_8-dihydrodiol-9_10-oxide						0.0003
Benzo[a]pyrene-7_8-diol			0.0013		0.0004	
Benzo[a]pyrene-7_8-oxide						0.0005
Benzo[a]pyrene-9_10-oxide						0.0005
beta-Alanyl-L-arginine				0.0003		
beta-Alanyl-L-lysine					0.0001	
Betaine		0.0003	0.0005			0.0005
Beta-tocopherol		0.0012	0.0013	0.0012	0.0012	
Bilirubin	0.0005	0.0024	0.0032	0.0012	0.0010	0.0008
Bilirubin glucuronide	0.0030		0.0031	0.0007		0.0008
Biliverdin			0.0006			
Biocytin	0.0009		0.0012		0.0014	0.0015
Biotin	0.0008	0.0011			0.0012	
Biotinyl-5^-AMP			0.0001			
Cadaverine			0.0004			
Caffeine	0.0002					
Calcidiol				0.0002	0.0008	0.0012
Capric acid						0.0004
Carnosine				0.0002		
CerP(d18:1/12:0)			0.0031		0.0014	
Chenodeoxycholic acid			0.0013			
Chenodeoxyglycocholic acid		0.0015	0.0019	0.0007	0.0008	
Chitin		0.0012				0.0030
Cholesterol				0.0001	0.0037	0.0007
Cholesterol sulfate					0.0009	0.0005
Choline	0.0003				0.0007	
Chondroitin			0.0007			
Citalopram aldehyde	0.0009			0.0010		
CMP-N-trimethyl-2-aminoethylphosphonate			0.0009			
Codeine-6-glucuronide	0.0004			0.0004		
Coproporphyrin III			0.0004			
Coproporphyrinogen I			0.0004			
Coproporphyrinogen III			0.0006			
Cortexolone			0.0000			
Corticosterone			0.0010			
Cytidine	0.0006					
D-4^-Phosphopantothenate					0.0005	
Debrisoquine			0.0002			
Dehydroepiandrosterone					0.0002	
Dehydroepiandrosterone sulfate				0.0001		
Deoxycorticosterone					0.0005	
Deoxycytidine					0.0005	0.0008
Deoxyguanosine		0.0013	0.0004		0.0012	
Deoxyinosine	0.0004				0.0010	
Deoxyribose 1-phosphate					0.0006	
Deoxyribose 5-phosphate					0.0004	
D-Erythrose 4-phosphate					0.0004	
DG(20:0/20:0/0:0)			0.0025			
D-Glutamic acid				0.0000		0.0001
D-Glutamine	0.0004					
Didemethylcitalopram				0.0005	0.0009	
Dihydrocortisol			0.0011			
Dihydrofolic acid						0.0020
Dihydrotestosterone	0.0014		0.0004			0.0007
Dihydrouracil		0.0000				
Dimethylglycine		0.0001				
D-Maltose					0.0013	
Docosa-4_7_10_13_16-pentaenoyl CoA		0.0016	0.0004			0.0046
Docosahexaenoic acid		0.0010	0.0004			
Dodecanoic acid			0.0004			
Dolichyl b-D-glucosyl phosphate			0.0001			
Dopamine	0.0006	0.0004	0.0000			
Dopamine 3-O-sulfate					0.0002	
D-Pantothenoyl-L-cysteine			0.0000	0.0012	0.0001	
D-Proline						0.0002
D-Ribose		0.0004				
D-Xylose		0.0003				
D-Xylulose		0.0001				
Eicosapentaenoic acid	0.0010	0.0014	0.0007		0.0016	
Elaidic acid		0.0018				0.0006
Enol-phenylpyruvate		0.0001	0.0000			
Estradiol					0.0007	
Estrone					0.0001	
Estrone glucuronide	0.0011	0.0020		0.0000	0.0017	0.0020
Estrone sulfate		0.0007	0.0004	0.0001		0.0002
Etiocholanedione					0.0007	
Etiocholanolone			0.0004			
Etiocholanolone glucuronide			0.0003			
Farnesyl pyrophosphate			0.0007			
Formiminoglutamic acid				0.0003		
Formylanthranilic acid				0.0001		
(Gal)2 (GlcA)1 (Xyl)1 (Ser)1			0.0001			
(Gal)2 (Xyl)1 (Ser)1				0.0003		
Gamma-Aminobutyric acid		0.0001				
Geranylgeranyl-PP			0.0008			
(GlcA)1 (GlcNAc)1				0.0001		
Gluconic acid		0.0005				
Gluconolactone					0.0003	
Glutarylcarnitine				0.0087		0.0015
Glyceric acid		0.0000				
Glycerol 3-phosphate						0.0001
Glycocholic acid	0.0007		0.0020	0.0016	0.0000	0.0022
Guanosine			0.0001	0.0001		
Guanosine monophosphate			0.0007			0.0010
Gulonic acid		0.0004				
Heptadecanoic acid		0.0005	0.0013			
Homocarnosine		0.0007			0.0004	
Homovanillic acid		0.0003	0.0001			
Indolepyruvate			0.0006			
L-3-Hydroxykynurenine					0.0004	
L-4-Hydroxyglutamate semialdehyde				0.0000		0.0001
L-Alpha-aminobutyric acid		0.0001				
L-Arabinose		0.0004				
L-Asparagine						0.0005
Lc4Cer				0.0005		
L-Cystathionine				0.0001		
L-Cystine		0.0002	0.0005			
Leukotriene A4			0.0004			
Leukotriene B4			0.0000			
Leukotriene C4	0.0028			0.0020	0.0013	0.0024
Leukotriene D4			0.0001			
L-Glutamic acid				0.0005		0.0005
L-Glutamine	0.0004					
L-Glyceric acid		0.0005	0.0004			
L-Gulonolactone					0.0002	
Linoleic acid		0.0012				
Liothyronine	0.0008					0.0004
L-Methionine				0.0005		
L-Octanoylcarnitine					0.0127	0.0013
L-Proline						0.0002
L-Threo-2-pentulose		0.0003				
L-Tryptophan		0.0003			0.0013	
L-Tyrosine		0.0005	0.0002			0.0006
L-Valine				0.0003	0.0005	0.0005
LysoPC(0:0/18:0)		0.0006	0.0002			0.0002
LysoSM(d18:1)		0.0007		0.0002		
Maleylacetoacetic acid			0.0003	0.0002		
Malonyl-Carnitin					0.0008	0.0005
Maltotriose		0.0023				
Menaquinol					0.0005	
Metanephrine					0.0007	
Mevalonic acid						0.0005
Myristic acid	0.0007	0.0012				
N10-Formyl-THF				0.0000	0.0019	0.0013
N(1)-(5-Phospho-D-ribosyl)glycinamide					0.0005	
N(2)-Formyl-N(1)-(5-phospho-D-ribosyl)glycinamide					0.0003	
N6_N6_N6-Trimethyl-L-lysine			0.0003		0.0004	
NA		0.0004	0.0004	0.0003	0.0003	
N-Acetylaspartylglutamylglutamate			0.0014			
N-Acetyl-D-Glucosamine 6-Phosphate			0.0004			
N-Acetyl-D-mannosamine 6-phosphate			0.0010			
N-Acetylneuraminate			0.0007		0.0007	
N-Acetylputrescine	0.0000					
NADH			0.0002			
Naphthalene-1_2-diol		0.0004				
Naphthalene epoxide		0.0004			0.0006	
Nervonic acid			0.0004			0.0008
Nicotinamide riboside			0.0001		0.0000	
Nicotinamide ribotide					0.0004	
Nicotinic acid adenine dinucleotide phosphate			0.0002			
N(omega)-Hydroxyarginine				0.0004	0.0006	
NPC			0.0009			
Obtusifoliol				0.0008		
Omeprazole	0.0000				0.0017	
OPC4-CoA					0.0001	
O-Phospho-4-hydroxy-L-threonine				0.0000		
Ophthalmic acid			0.0000			
Orotidylic acid					0.0010	
Ortho-Hydroxyphenylacetic acid			0.0000			
Oxalacetic acid						0.0001
Oxidized glutathione		0.0025			0.0025	
Oxoadipic acid		0.0005				
Oxoglutaric acid				0.0002	0.0003	
Palmitic acid			0.0004			
Pantothenic acid	0.0003		0.0010		0.0004	
Pentadecanoic acid			0.0009			
Pentaglutamyl folate	0.0042				0.0037	
Phenylacetic acid		0.0002	0.0001			
Phenylethylamine		0.0001				
Phenylpyruvic acid		0.0001	0.0000			
p-Hydroxyphenylacetic acid			0.0002			
Pregnenolone		0.0004				
Presqualene diphosphate			0.0004			
Propinol adenylate			0.0002			
Prostaglandin D2	0.0018		0.0004	0.0009		0.0008
Prostaglandin E2	0.0018		0.0004	0.0009		0.0008
Prostaglandin F2a			0.0006	0.0004		0.0003
Prostaglandin H2			0.0000	0.0004		
Prostaglandin I2			0.0007	0.0000		
Protoporphyrin IX			0.0009	0.0004		
Protoporphyrinogen IX			0.0002			
Putrescine			0.0001			
Pyridoxamine		0.0006				
Pyroglutamic acid				0.0001		0.0004
Pyrrolidonecarboxylic acid				0.0001		0.0004
Pyrroline hydroxycarboxylic acid				0.0002		0.0004
(R)-3-Ureidoisobutyrate	0.0004					
(R)-b-aminoisobutyric acid	0.0001	0.0001	0.0004			
Retinal		0.0003				
Retinoyl b-glucuronide	0.0001	0.0003	0.0020			
Riboflavin	0.0000				0.0000	
Riboflavin reduced			0.0006			
S-Adenosylhomocysteine					0.0000	
(S)-b-aminoisobutyric acid	0.0001	0.0001	0.0004			
Selenomethionine					0.0003	
Serotonin			0.0003			
SN-38					0.0001	
SN38 glucuronide			0.0005			
Sphinganine 1-phosphate	0.0002		0.0019		0.0019	
Sphingosine						0.0001
Sphingosine 1-phosphate		0.0008		0.0000		
Squalene				0.0004		
Stachyose			0.0000			
Stearidonic acid			0.0006		0.0008	
Sucrose					0.0013	
Tamoxifen N-oxide				0.0016		
T antigen			0.0006		0.0002	
Taurochenodesoxycholic acid	0.0011	0.0018	0.0021	0.0016	0.0009	0.0006
Taurocholic acid	0.0013	0.0019		0.0021		0.0010
Testosterone					0.0003	
Tetracosahexaenoic acid				0.0008	0.0001	0.0010
Tetracosanoic acid			0.0007			
Tetrahydrocortisol				0.0001		
Tetrahydrofolic acid	0.0009		0.0001	0.0005	0.0012	0.0019
Tetrahydrofolyl-[Glu](n)			0.0012			
TG(8:0/8:0/8:0)	0.0021		0.0009	0.0001		0.0025
THF-L-glutamate			0.0003			
Thiamine						0.0008
Thiamine monophosphate						0.0014
Thiamine pyrophosphate				0.0005		
Thromboxane A2	0.0018		0.0004	0.0009		0.0008
Thymidine					0.0004	
Thyroxine		0.0022	0.0025			
trans-3_4-Dihydro-3_4-dihydroxy-7_12-dimethylbenz[a]anthracene			0.0002			
trans-5_6-Dihydro-5_6-dihydroxy-7_12-dimethylbenz[a]anthracene			0.0002			
Trehalose					0.0013	
Triiodothyronine sulfate			0.0014			
Trypanothione			0.0017			
Tyramine-O-sulfate	0.0001				0.0005	
Uracil			0.0006			
Ureidopropionic acid						0.0005
Uridine	0.0005	0.0007	0.0009	0.0000	0.0006	
Uridine 5^-monophosphate					0.0008	
Uridine triphosphate	0.0014					
Urobilinogen	0.0017		0.0000	0.0003		
Vaccenic acid		0.0018				0.0006
Vitamin A	0.0010		0.0003			
Vitamin D3		0.0016	0.0022	0.0000		0.0010
Xanthosine					0.0000	
(Xyl)1 (Ser)1						0.0003
MS_METABOLITE_DATA_END
#METABOLITES
METABOLITES_START
metabolite_name	pubchem_id	inchi_key	kegg_id	other_id	other_id_type	ri	ri_type	moverz_quant	
11,12,15-THETA									
11,14,15-THETA									
11b,17a,21-Trihydroxypreg-nenolone									
11b,21-Dihydroxy-3,20-oxo-5b-pregnan-18-al									
11-cis-Retinaldehyde									
11-cis-Retinol									
11H-14,15-EETA									
1,2-Dihydronaphthalene-1,2-diol									
12(R)-HPETE									
12(S)-HPETE									
15H-11,12-EETA									
15(S)-HPETE									
16a-Hydroxyandrost-4-ene-3,17-dione									
16b-Hydroxyestrone									
17alpha,21-Dihydroxypregnenolone									
17-beta-Estradiol-3-glucuronide									
17-Hydroxyprogesterone									
19-Hydroxyandrost-4-ene-3,17-dione									
19-Oxoandrost-4-ene-3,17-dione									
19-Oxotestosterone									
1a,11b-Dihydro-4,9-dimethylbenz[a]anthra[3,4-b]oxirene									
(1aalpha,2beta,3alpha,11calpha)-1a,2,3,11c-Tetrahydro-6,11-dimethylbenzo[6,7]phenanthro[3,4-b]oxirene-2,3-diol									
1-Methylxanthine									
1-Naphthol									
1-Nitronaphthalene-5,6-oxide									
1-Nitronaphthalene-7,8-oxide									
1-Pyrroline-4-hydroxy-2-carboxylate									
20a-Dihydroprogesterone									
20-Hydroxy-leukotriene B4									
21-Deoxycortisol									
24-Hydroxycalcitriol									
2-Hydroxy-3-(4-hydroxyphenyl)propenoic acid									
2-Hydroxyestrone									
2-Methoxyestradiol									
2-Methoxy-estradiol-17b 3-glucuronide									
2-Methoxyestrone									
2-Oxo-4-methylthiobutanoic acid									
2-Oxoarginine									
2-(S-Glutathionyl)acetyl glutathione									
3,4-Dihydroxyphenylacetaldehyde									
3,4-Dihydroxyphenylglycol									
3a,21-Dihydroxy-5b-pregnane-11,20-dione									
3a,7a,12a-Trihydroxy-5b-cholestan-26-al									
3a,7a-Dihydroxycoprostanic acid									
3beta,7alpha-Dihydroxy-5-cholestenoate									
3beta-Hydroxypregn-5-en-20-one sulfate									
3-Hydroxylidocaine									
3-ketosphinganine									
3-Methoxy-4-hydroxyphenylglycolaldehyde									
4a-Carboxy-4b-methyl-5a-cholesta-8,24-dien-3b-ol									
4a-Hydroxytetrahydrobiopterin									
4-Guanidinobutanoic acid									
4-Hydroxy-4-(methylnitrosoamino)-1-(3-pyridinyl)-1-butanone									
4-Hydroxydebrisoquine									
4-[(Hydroxymethyl)nitrosoamino]-1-(3-pyridinyl)-1-butanone									
4-Hydroxyphenylacetaldehyde									
4-Hydroxyphenylpyruvic acid									
4-Hydroxy tolbutamide									
4-Pyridoxic acid									
4-Trimethylammoniobutanoic acid									
5,10-Methylene-THF									
5,6-Dihydro-5-fluorouracil									
5,7,24(28)-Ergostatrienol									
5-Acetylamino-6-formylamino-3-methyluracil									
5a-Dihydrotestosterone sulfate									
5-alpha-Dihydrotestosterone glucuronide									
5-Androstenediol									
5a-Pregnane-3,20-dione									
5b-Dihydrotestosterone									
5-Fluorouridine									
5-Formiminotetrahydrofolic acid									
5-Hydroxykynurenine									
5-L-Glutamyl-taurine									
5-Methoxyindoleacetate									
5-Methyltetrahydrofolic acid									
5'-Methylthioadenosine									
5-phosphonooxy-L-lysine									
5(S)-Hydroperoxyeicosatetraenoic acid									
6-Hydroxymelatonin									
6-Methylthiopurine									
7,12-Dimethylbenz[a]anthracene 5,6-oxide									
7,8-Dihydroneopterin									
7alpha-Hydroxy-3-oxo-4-cholestenoate									
7-Hydroxymethyl-12-methylbenz[a]anthracene									
7-Hydroxymethyl-12-methylbenz[a]anthracene sulfate									
7-Methylxanthine									
8-Isoprostaglandin F2a									
8(S)-HPETE									
9-cis-Retinal									
9-cis-Retinoic acid									
9-cis-Retinol									
9-Hydroxybenzo[a]pyrene									
Acetylcholine chloride									
Acetyl-CoA									
Adenosine									
ADP-ribose 2'-phosphate									
Adrenic acid									
Adrenosterone									
Aflatoxin B1									
all-trans-Decaprenyl diphosphate									
All-trans-retinoic acid									
alpha-Fluoro-beta-ureidopropionic acid									
Alpha-Lactose									
Alpha-Tocotrienol									
Aminoadipic acid									
Androstanedione									
Androstenedione									
Androsterone									
Androsterone glucuronide									
Anserine									
Arachidic acid									
Arachidonic acid									
Argininosuccinic acid									
B-Carotene									
Benzo[a]pyrene									
Benzo[a]pyrene-4,5-oxide									
Benzo[a]pyrene-7,8-dihydrodiol-9,10-oxide									
Benzo[a]pyrene-7,8-diol									
Benzo[a]pyrene-7,8-oxide									
Benzo[a]pyrene-9,10-oxide									
beta-Alanyl-L-arginine									
beta-Alanyl-L-lysine									
Betaine									
Beta-tocopherol									
Bilirubin									
Bilirubin glucuronide									
Biliverdin									
Biocytin									
Biotin									
Biotinyl-5'-AMP									
Cadaverine									
Caffeine									
Calcidiol									
Capric acid									
Carnosine									
CerP(d18:1/12:0)									
Chenodeoxycholic acid									
Chenodeoxyglycocholic acid									
Chitin									
Cholesterol									
Cholesterol sulfate									
Choline									
Chondroitin									
Citalopram aldehyde									
CMP-N-trimethyl-2-aminoethylphosphonate									
Codeine-6-glucuronide									
Coproporphyrin III									
Coproporphyrinogen I									
Coproporphyrinogen III									
Cortexolone									
Corticosterone									
Cytidine									
D-4'-Phosphopantothenate									
Debrisoquine									
Dehydroepiandrosterone									
Dehydroepiandrosterone sulfate									
Deoxycorticosterone									
Deoxycytidine									
Deoxyguanosine									
Deoxyinosine									
Deoxyribose 1-phosphate									
Deoxyribose 5-phosphate									
D-Erythrose 4-phosphate									
DG(20:0/20:0/0:0)									
D-Glutamic acid									
D-Glutamine									
Didemethylcitalopram									
Dihydrocortisol									
Dihydrofolic acid									
Dihydrotestosterone									
Dihydrouracil									
Dimethylglycine									
D-Maltose									
Docosa-4,7,10,13,16-pentaenoyl CoA									
Docosahexaenoic acid									
Dodecanoic acid									
Dolichyl b-D-glucosyl phosphate									
Dopamine									
Dopamine 3-O-sulfate									
D-Pantothenoyl-L-cysteine									
D-Proline									
D-Ribose									
D-Xylose									
D-Xylulose									
Eicosapentaenoic acid									
Elaidic acid									
Enol-phenylpyruvate									
Estradiol									
Estrone									
Estrone glucuronide									
Estrone sulfate									
Etiocholanedione									
Etiocholanolone									
Etiocholanolone glucuronide									
Farnesyl pyrophosphate									
Formiminoglutamic acid									
Formylanthranilic acid									
(Gal)2 (GlcA)1 (Xyl)1 (Ser)1									
(Gal)2 (Xyl)1 (Ser)1									
Gamma-Aminobutyric acid									
Geranylgeranyl-PP									
(GlcA)1 (GlcNAc)1									
Gluconic acid									
Gluconolactone									
Glutarylcarnitine									
Glyceric acid									
Glycerol 3-phosphate									
Glycocholic acid									
Guanosine									
Guanosine monophosphate									
Gulonic acid									
Heptadecanoic acid									
Homocarnosine									
Homovanillic acid									
Indolepyruvate									
L-3-Hydroxykynurenine									
L-4-Hydroxyglutamate semialdehyde									
L-Alpha-aminobutyric acid									
L-Arabinose									
L-Asparagine									
Lc4Cer									
L-Cystathionine									
L-Cystine									
Leukotriene A4									
Leukotriene B4									
Leukotriene C4									
Leukotriene D4									
L-Glutamic acid									
L-Glutamine									
L-Glyceric acid									
L-Gulonolactone									
Linoleic acid									
Liothyronine									
L-Methionine									
L-Octanoylcarnitine									
L-Proline									
L-Threo-2-pentulose									
L-Tryptophan									
L-Tyrosine									
L-Valine									
LysoPC(0:0/18:0)									
LysoSM(d18:1)									
Maleylacetoacetic acid									
Malonyl-Carnitin									
Maltotriose									
Menaquinol									
Metanephrine									
Mevalonic acid									
Myristic acid									
N10-Formyl-THF									
N(1)-(5-Phospho-D-ribosyl)glycinamide									
N(2)-Formyl-N(1)-(5-phospho-D-ribosyl)glycinamide									
N6,N6,N6-Trimethyl-L-lysine									
NA									
N-Acetylaspartylglutamylglutamate									
N-Acetyl-D-Glucosamine 6-Phosphate									
N-Acetyl-D-mannosamine 6-phosphate									
N-Acetylneuraminate									
N-Acetylputrescine									
NADH									
Naphthalene-1,2-diol									
Naphthalene epoxide									
Nervonic acid									
Nicotinamide riboside									
Nicotinamide ribotide									
Nicotinic acid adenine dinucleotide phosphate									
N(omega)-Hydroxyarginine									
NPC									
Obtusifoliol									
Omeprazole									
OPC4-CoA									
O-Phospho-4-hydroxy-L-threonine									
Ophthalmic acid									
Orotidylic acid									
Ortho-Hydroxyphenylacetic acid									
Oxalacetic acid									
Oxidized glutathione									
Oxoadipic acid									
Oxoglutaric acid									
Palmitic acid									
Pantothenic acid									
Pentadecanoic acid									
Pentaglutamyl folate									
Phenylacetic acid									
Phenylethylamine									
Phenylpyruvic acid									
p-Hydroxyphenylacetic acid									
Pregnenolone									
Presqualene diphosphate									
Propinol adenylate									
Prostaglandin D2									
Prostaglandin E2									
Prostaglandin F2a									
Prostaglandin H2									
Prostaglandin I2									
Protoporphyrin IX									
Protoporphyrinogen IX									
Putrescine									
Pyridoxamine									
Pyroglutamic acid									
Pyrrolidonecarboxylic acid									
Pyrroline hydroxycarboxylic acid									
(R)-3-Ureidoisobutyrate									
(R)-b-aminoisobutyric acid									
Retinal									
Retinoyl b-glucuronide									
Riboflavin									
Riboflavin reduced									
S-Adenosylhomocysteine									
(S)-b-aminoisobutyric acid									
Selenomethionine									
Serotonin									
SN-38									
SN38 glucuronide									
Sphinganine 1-phosphate									
Sphingosine									
Sphingosine 1-phosphate									
Squalene									
Stachyose									
Stearidonic acid									
Sucrose									
Tamoxifen N-oxide									
T antigen									
Taurochenodesoxycholic acid									
Taurocholic acid									
Testosterone									
Tetracosahexaenoic acid									
Tetracosanoic acid									
Tetrahydrocortisol									
Tetrahydrofolic acid									
Tetrahydrofolyl-[Glu](n)									
TG(8:0/8:0/8:0)									
THF-L-glutamate									
Thiamine									
Thiamine monophosphate									
Thiamine pyrophosphate									
Thromboxane A2									
Thymidine									
Thyroxine									
trans-3,4-Dihydro-3,4-dihydroxy-7,12-dimethylbenz[a]anthracene									
trans-5,6-Dihydro-5,6-dihydroxy-7,12-dimethylbenz[a]anthracene									
Trehalose									
Triiodothyronine sulfate									
Trypanothione									
Tyramine-O-sulfate									
Uracil									
Ureidopropionic acid									
Uridine									
Uridine 5'-monophosphate									
Uridine triphosphate									
Urobilinogen									
Vaccenic acid									
Vitamin A									
Vitamin D3									
Xanthosine									
(Xyl)1 (Ser)1									
METABOLITES_END
#END