{
"METABOLOMICS WORKBENCH":{"STUDY_ID":"ST001411","ANALYSIS_ID":"AN002361","VERSION":"1","CREATED_ON":"June 23, 2020, 8:25 am"},

"PROJECT":{"PROJECT_TITLE":"Plasma metabolites of lipid metabolism associate with diabetic polyneuropathy in a cohort with screen-tested type 2 diabetes: ADDITION-Denmark","PROJECT_SUMMARY":"The global rise in type 2 diabetes (T2D) is associated with a concomitant increase in diabetic complications. Diabetic polyneuropathy (DPN), the most frequent T2D complication, is characterized by sensory peripheral nerve damage. Although managing glucose effectively slows DPN progression in type 1 diabetes patients, it has limited efficacy in neuropathic T2D patients. The metabolic syndrome (MetS) recently emerged as a major risk factor for DPN; however, the metabolites associated with MetS that correlate with DPN are unknown. We conducted a global plasma metabolomics analysis from a cohort of patients enrolled in the Anglo-Danish-Dutch study of Intensive Treatment of Diabetes in Primary Care (ADDITION), including healthy control subjects, T2D patients, and T2D DPN patients. We identified 15 total plasma metabolites that were altered in T2D DPN patients, including lipids, amino acids, and energy-related metabolites. We evaluated the correlation between these metabolites and all lipid species to identify major changes in both plasma free fatty acids and complex lipids in T2D DPN patients, and found significant alterations in the abundance of long-chain saturated fatty acids, acylcarnitines, and sphingolipids. Our study suggests that DPN in T2D is associated with novel alterations in plasma metabolites related to lipid metabolism.","INSTITUTE":"University of Michigan","DEPARTMENT":"Neurology Department","LABORATORY":"Feldman's lab","LAST_NAME":"Feldman","FIRST_NAME":"Eva","ADDRESS":"5017 AATBSRB, 109 Zina Pitcher Place, Ann Arbor, MI 48109-2200","EMAIL":"efeldman@med.umich.edu","PHONE":"7347637274","FUNDING_SOURCE":"Support for this research was provided by the Novo Nordisk Foundation through a Novo Nordisk Foundation Challenge Programme grant (NNF14OC0011633), the National Institutes of Health (1R24082841, 1R21NS102924, and 1DP3DK094292), the National Institute for Diabetes and Digestive and Kidney Diseases (NIDDK) (1F32DK112642 and 1K99DK119366), the American Diabetes Association (7-12-BS-045), the NeuroNetwork for Emerging Therapies, and the A. Alfred Taubman Medical Research Institute."},

"STUDY":{"STUDY_TITLE":"Plasma metabolites of lipid metabolism associate with diabetic polyneuropathy in a cohort with screen-tested type 2 diabetes: ADDITION-Denmark","STUDY_SUMMARY":"The global rise in type 2 diabetes (T2D) is associated with a concomitant increase in diabetic complications. Diabetic polyneuropathy (DPN), the most frequent T2D complication, is characterized by sensory peripheral nerve damage. Although managing glucose effectively slows DPN progression in type 1 diabetes patients, it has limited efficacy in neuropathic T2D patients. The metabolic syndrome (MetS) recently emerged as a major risk factor for DPN; however, the metabolites associated with MetS that correlate with DPN are unknown. We conducted a global plasma metabolomics analysis from a cohort of patients enrolled in the Anglo-Danish-Dutch study of Intensive Treatment of Diabetes in Primary Care (ADDITION), including healthy control subjects, T2D patients, and T2D DPN patients. We identified 15 total plasma metabolites that were altered in T2D DPN patients, including lipids, amino acids, and energy-related metabolites. We evaluated the correlation between these metabolites and all lipid species to identify major changes in both plasma free fatty acids and complex lipids in T2D DPN patients, and found significant alterations in the abundance of long-chain saturated fatty acids, acylcarnitines, and sphingolipids. Our study suggests that DPN in T2D is associated with novel alterations in plasma metabolites related to lipid metabolism.","INSTITUTE":"University of Michigan","LAST_NAME":"Feldman","FIRST_NAME":"Eva","ADDRESS":"5017 AATBSRB, 109 Zina Pitcher Place, Ann Arbor, MI 48109-2200","EMAIL":"efeldman@med.umich.edu","PHONE":"7347637274","NUM_GROUPS":"3","TOTAL_SUBJECTS":"106","NUM_MALES":"83","NUM_FEMALES":"23"},

"SUBJECT":{"SUBJECT_TYPE":"Human","SUBJECT_SPECIES":"Homo sapiens","TAXONOMY_ID":"9606"},
"SUBJECT_SAMPLE_FACTORS":[
{
"Subject ID":"-",
"Sample ID":"MICH-01419",
"Factors":{"Group":"Diabetic Neuropathy"}
},
{
"Subject ID":"-",
"Sample ID":"MICH-01420",
"Factors":{"Group":"Diabetic non neuropathy"}
},
{
"Subject ID":"-",
"Sample ID":"MICH-01421",
"Factors":{"Group":"Diabetic non neuropathy"}
},
{
"Subject ID":"-",
"Sample ID":"MICH-01422",
"Factors":{"Group":"Diabetic Neuropathy"}
},
{
"Subject ID":"-",
"Sample ID":"MICH-01423",
"Factors":{"Group":"Diabetic non neuropathy"}
},
{
"Subject ID":"-",
"Sample ID":"MICH-01424",
"Factors":{"Group":"Diabetic non neuropathy"}
},
{
"Subject ID":"-",
"Sample ID":"MICH-01425",
"Factors":{"Group":"Diabetic Neuropathy"}
},
{
"Subject ID":"-",
"Sample ID":"MICH-01426",
"Factors":{"Group":"Diabetic Neuropathy"}
},
{
"Subject ID":"-",
"Sample ID":"MICH-01427",
"Factors":{"Group":"Diabetic non neuropathy"}
},
{
"Subject ID":"-",
"Sample ID":"MICH-01428",
"Factors":{"Group":"Diabetic Neuropathy"}
},
{
"Subject ID":"-",
"Sample ID":"MICH-01429",
"Factors":{"Group":"Diabetic Neuropathy"}
},
{
"Subject ID":"-",
"Sample ID":"MICH-01430",
"Factors":{"Group":"Diabetic Neuropathy"}
},
{
"Subject ID":"-",
"Sample ID":"MICH-01431",
"Factors":{"Group":"Diabetic Neuropathy"}
},
{
"Subject ID":"-",
"Sample ID":"MICH-01432",
"Factors":{"Group":"Diabetic Neuropathy"}
},
{
"Subject ID":"-",
"Sample ID":"MICH-01433",
"Factors":{"Group":"Diabetic non neuropathy"}
},
{
"Subject ID":"-",
"Sample ID":"MICH-01434",
"Factors":{"Group":"Diabetic Neuropathy"}
},
{
"Subject ID":"-",
"Sample ID":"MICH-01435",
"Factors":{"Group":"Diabetic non neuropathy"}
},
{
"Subject ID":"-",
"Sample ID":"MICH-01436",
"Factors":{"Group":"Diabetic Neuropathy"}
},
{
"Subject ID":"-",
"Sample ID":"MICH-01437",
"Factors":{"Group":"Diabetic non neuropathy"}
},
{
"Subject ID":"-",
"Sample ID":"MICH-01438",
"Factors":{"Group":"Diabetic non neuropathy"}
},
{
"Subject ID":"-",
"Sample ID":"MICH-01439",
"Factors":{"Group":"Diabetic Neuropathy"}
},
{
"Subject ID":"-",
"Sample ID":"MICH-01440",
"Factors":{"Group":"Diabetic non neuropathy"}
},
{
"Subject ID":"-",
"Sample ID":"MICH-01441",
"Factors":{"Group":"Diabetic Neuropathy"}
},
{
"Subject ID":"-",
"Sample ID":"MICH-01442",
"Factors":{"Group":"Diabetic non neuropathy"}
},
{
"Subject ID":"-",
"Sample ID":"MICH-01443",
"Factors":{"Group":"Diabetic non neuropathy"}
},
{
"Subject ID":"-",
"Sample ID":"MICH-01444",
"Factors":{"Group":"Diabetic Neuropathy"}
},
{
"Subject ID":"-",
"Sample ID":"MICH-01445",
"Factors":{"Group":"Diabetic non neuropathy"}
},
{
"Subject ID":"-",
"Sample ID":"MICH-01446",
"Factors":{"Group":"Diabetic Neuropathy"}
},
{
"Subject ID":"-",
"Sample ID":"MICH-01447",
"Factors":{"Group":"Diabetic Neuropathy"}
},
{
"Subject ID":"-",
"Sample ID":"MICH-01448",
"Factors":{"Group":"Diabetic non neuropathy"}
},
{
"Subject ID":"-",
"Sample ID":"MICH-01449",
"Factors":{"Group":"Diabetic non neuropathy"}
},
{
"Subject ID":"-",
"Sample ID":"MICH-01450",
"Factors":{"Group":"Diabetic non neuropathy"}
},
{
"Subject ID":"-",
"Sample ID":"MICH-01451",
"Factors":{"Group":"Diabetic non neuropathy"}
},
{
"Subject ID":"-",
"Sample ID":"MICH-01452",
"Factors":{"Group":"Diabetic Neuropathy"}
},
{
"Subject ID":"-",
"Sample ID":"MICH-01453",
"Factors":{"Group":"Diabetic Neuropathy"}
},
{
"Subject ID":"-",
"Sample ID":"MICH-01454",
"Factors":{"Group":"Diabetic Neuropathy"}
},
{
"Subject ID":"-",
"Sample ID":"MICH-01455",
"Factors":{"Group":"Diabetic Neuropathy"}
},
{
"Subject ID":"-",
"Sample ID":"MICH-01456",
"Factors":{"Group":"Diabetic non neuropathy"}
},
{
"Subject ID":"-",
"Sample ID":"MICH-01457",
"Factors":{"Group":"Diabetic non neuropathy"}
},
{
"Subject ID":"-",
"Sample ID":"MICH-01458",
"Factors":{"Group":"Diabetic non neuropathy"}
},
{
"Subject ID":"-",
"Sample ID":"MICH-01459",
"Factors":{"Group":"Diabetic Neuropathy"}
},
{
"Subject ID":"-",
"Sample ID":"MICH-01460",
"Factors":{"Group":"Diabetic non neuropathy"}
},
{
"Subject ID":"-",
"Sample ID":"MICH-01461",
"Factors":{"Group":"Diabetic non neuropathy"}
},
{
"Subject ID":"-",
"Sample ID":"MICH-01462",
"Factors":{"Group":"Diabetic Neuropathy"}
},
{
"Subject ID":"-",
"Sample ID":"MICH-01463",
"Factors":{"Group":"Diabetic Neuropathy"}
},
{
"Subject ID":"-",
"Sample ID":"MICH-01464",
"Factors":{"Group":"Diabetic Neuropathy"}
},
{
"Subject ID":"-",
"Sample ID":"MICH-01465",
"Factors":{"Group":"Diabetic Neuropathy"}
},
{
"Subject ID":"-",
"Sample ID":"MICH-01466",
"Factors":{"Group":"Diabetic Neuropathy"}
},
{
"Subject ID":"-",
"Sample ID":"MICH-01467",
"Factors":{"Group":"Diabetic non neuropathy"}
},
{
"Subject ID":"-",
"Sample ID":"MICH-01468",
"Factors":{"Group":"Diabetic Neuropathy"}
},
{
"Subject ID":"-",
"Sample ID":"MICH-01469",
"Factors":{"Group":"Diabetic Neuropathy"}
},
{
"Subject ID":"-",
"Sample ID":"MICH-01470",
"Factors":{"Group":"Diabetic Neuropathy"}
},
{
"Subject ID":"-",
"Sample ID":"MICH-01471",
"Factors":{"Group":"Diabetic non neuropathy"}
},
{
"Subject ID":"-",
"Sample ID":"MICH-01472",
"Factors":{"Group":"Diabetic Neuropathy"}
},
{
"Subject ID":"-",
"Sample ID":"MICH-01473",
"Factors":{"Group":"Diabetic Neuropathy"}
},
{
"Subject ID":"-",
"Sample ID":"MICH-01474",
"Factors":{"Group":"Diabetic non neuropathy"}
},
{
"Subject ID":"-",
"Sample ID":"MICH-01475",
"Factors":{"Group":"Diabetic non neuropathy"}
},
{
"Subject ID":"-",
"Sample ID":"MICH-01476",
"Factors":{"Group":"Diabetic non neuropathy"}
},
{
"Subject ID":"-",
"Sample ID":"MICH-01477",
"Factors":{"Group":"Diabetic non neuropathy"}
},
{
"Subject ID":"-",
"Sample ID":"MICH-01478",
"Factors":{"Group":"Diabetic non neuropathy"}
},
{
"Subject ID":"-",
"Sample ID":"MICH-01479",
"Factors":{"Group":"Diabetic non neuropathy"}
},
{
"Subject ID":"-",
"Sample ID":"MICH-01480",
"Factors":{"Group":"Diabetic non neuropathy"}
},
{
"Subject ID":"-",
"Sample ID":"MICH-01481",
"Factors":{"Group":"Diabetic Neuropathy"}
},
{
"Subject ID":"-",
"Sample ID":"MICH-01482",
"Factors":{"Group":"Diabetic Neuropathy"}
},
{
"Subject ID":"-",
"Sample ID":"MICH-01483",
"Factors":{"Group":"Diabetic Neuropathy"}
},
{
"Subject ID":"-",
"Sample ID":"MICH-01484",
"Factors":{"Group":"Diabetic non neuropathy"}
},
{
"Subject ID":"-",
"Sample ID":"MICH-01485",
"Factors":{"Group":"Diabetic Neuropathy"}
},
{
"Subject ID":"-",
"Sample ID":"MICH-01486",
"Factors":{"Group":"Diabetic non neuropathy"}
},
{
"Subject ID":"-",
"Sample ID":"MICH-01487",
"Factors":{"Group":"Diabetic non neuropathy"}
},
{
"Subject ID":"-",
"Sample ID":"MICH-01488",
"Factors":{"Group":"Diabetic non neuropathy"}
},
{
"Subject ID":"-",
"Sample ID":"MICH-01489",
"Factors":{"Group":"Diabetic non neuropathy"}
},
{
"Subject ID":"-",
"Sample ID":"MICH-01490",
"Factors":{"Group":"Diabetic Neuropathy"}
},
{
"Subject ID":"-",
"Sample ID":"MICH-01491",
"Factors":{"Group":"Diabetic Neuropathy"}
},
{
"Subject ID":"-",
"Sample ID":"MICH-01492",
"Factors":{"Group":"Diabetic Neuropathy"}
},
{
"Subject ID":"-",
"Sample ID":"MICH-01493",
"Factors":{"Group":"Diabetic non neuropathy"}
},
{
"Subject ID":"-",
"Sample ID":"MICH-01494",
"Factors":{"Group":"Diabetic non neuropathy"}
},
{
"Subject ID":"-",
"Sample ID":"MICH-01495",
"Factors":{"Group":"Diabetic Neuropathy"}
},
{
"Subject ID":"-",
"Sample ID":"MICH-01496",
"Factors":{"Group":"Diabetic Neuropathy"}
},
{
"Subject ID":"-",
"Sample ID":"MICH-01497",
"Factors":{"Group":"Diabetic non neuropathy"}
},
{
"Subject ID":"-",
"Sample ID":"MICH-01498",
"Factors":{"Group":"Diabetic non neuropathy"}
},
{
"Subject ID":"-",
"Sample ID":"MICH-01499",
"Factors":{"Group":"Diabetic non neuropathy"}
},
{
"Subject ID":"-",
"Sample ID":"MICH-01500",
"Factors":{"Group":"Diabetic non neuropathy"}
},
{
"Subject ID":"-",
"Sample ID":"MICH-01501",
"Factors":{"Group":"Diabetic non neuropathy"}
},
{
"Subject ID":"-",
"Sample ID":"MICH-01502",
"Factors":{"Group":"Diabetic Neuropathy"}
},
{
"Subject ID":"-",
"Sample ID":"MICH-01503",
"Factors":{"Group":"Diabetic non neuropathy"}
},
{
"Subject ID":"-",
"Sample ID":"MICH-01504",
"Factors":{"Group":"Diabetic Neuropathy"}
},
{
"Subject ID":"-",
"Sample ID":"MICH-01505",
"Factors":{"Group":"Diabetic non neuropathy"}
},
{
"Subject ID":"-",
"Sample ID":"MICH-01506",
"Factors":{"Group":"Diabetic non neuropathy"}
},
{
"Subject ID":"-",
"Sample ID":"MICH-01507",
"Factors":{"Group":"Diabetic Neuropathy"}
},
{
"Subject ID":"-",
"Sample ID":"MICH-01508",
"Factors":{"Group":"Diabetic non neuropathy"}
},
{
"Subject ID":"-",
"Sample ID":"MICH-01509",
"Factors":{"Group":"Diabetic non neuropathy"}
},
{
"Subject ID":"-",
"Sample ID":"MICH-01510",
"Factors":{"Group":"Diabetic Neuropathy"}
},
{
"Subject ID":"-",
"Sample ID":"MICH-01511",
"Factors":{"Group":"Diabetic Neuropathy"}
},
{
"Subject ID":"-",
"Sample ID":"MICH-01512",
"Factors":{"Group":"Diabetic Neuropathy"}
},
{
"Subject ID":"-",
"Sample ID":"MICH-01513",
"Factors":{"Group":"Diabetic non neuropathy"}
},
{
"Subject ID":"-",
"Sample ID":"MICH-01514",
"Factors":{"Group":"Diabetic Neuropathy"}
},
{
"Subject ID":"-",
"Sample ID":"MICH-01515",
"Factors":{"Group":"Diabetic Neuropathy"}
},
{
"Subject ID":"-",
"Sample ID":"MICH-01516",
"Factors":{"Group":"Normal"}
},
{
"Subject ID":"-",
"Sample ID":"MICH-01517",
"Factors":{"Group":"Normal"}
},
{
"Subject ID":"-",
"Sample ID":"MICH-01518",
"Factors":{"Group":"Normal"}
},
{
"Subject ID":"-",
"Sample ID":"MICH-01519",
"Factors":{"Group":"Normal"}
},
{
"Subject ID":"-",
"Sample ID":"MICH-01520",
"Factors":{"Group":"Normal"}
},
{
"Subject ID":"-",
"Sample ID":"MICH-01521",
"Factors":{"Group":"Normal"}
},
{
"Subject ID":"-",
"Sample ID":"MICH-01522",
"Factors":{"Group":"Normal"}
},
{
"Subject ID":"-",
"Sample ID":"MICH-01523",
"Factors":{"Group":"Normal"}
},
{
"Subject ID":"-",
"Sample ID":"MICH-01524",
"Factors":{"Group":"Normal"}
}
],
"COLLECTION":{"COLLECTION_SUMMARY":"At the mean 13 year follow up after T2D diagnosis, blood samples were collected from patients the same day as anthropometrics and DPN assessment. Plasma was collected in purple EDTA tubes with 10 mL of buffy coat, inverted 10 times, incubated for 30-90 minutes at room temperature, and centrifuged at 3000 rpm for 10 minutes. The plasma supernatant was collected by aspirating plasma to approximately 5 mm above the buffy coat and plasma samples were stored in 0.5 mL aliquots at -80C prior to metabolomics analysis.","SAMPLE_TYPE":"Blood (plasma)"},

"TREATMENT":{"TREATMENT_SUMMARY":"NA"},

"SAMPLEPREP":{"SAMPLEPREP_SUMMARY":"Samples were prepared using the automated MicroLab STAR® system from Hamilton Company. Several recovery standards were added prior to the first step in the extraction process for QC purposes. To remove protein, dissociate small molecules bound to protein or trapped in the precipitated protein matrix, and to recover chemically diverse metabolites, proteins were precipitated with methanol under vigorous shaking for 2 min (Glen Mills GenoGrinder 2000) followed by centrifugation. The resulting extract was divided into five fractions: two for analysis by two separate reverse phase (RP)/UPLC-MS/MS methods with positive ion mode electrospray ionization (ESI), one for analysis by RP/UPLC-MS/MS with negative ion mode ESI, one for analysis by HILIC/UPLC-MS/MS with negative ion mode ESI, and one sample was reserved for backup. Samples were placed briefly on a TurboVap® (Zymark) to remove the organic solvent. The sample extracts were stored overnight under nitrogen before preparation for analysis."},

"CHROMATOGRAPHY":{"CHROMATOGRAPHY_TYPE":"HILIC","INSTRUMENT_NAME":"Waters Acquity","COLUMN_NAME":"Waters Acquity CSH C18 (100 x 2.1mm, 1.7um)"},

"ANALYSIS":{"ANALYSIS_TYPE":"MS","LABORATORY_NAME":"Metabolon"},

"MS":{"INSTRUMENT_NAME":"Waters Acquity","INSTRUMENT_TYPE":"UPLC-MS","MS_TYPE":"ESI","ION_MODE":"POSITIVE","MS_COMMENTS":"The informatics system consisted of four major components, the Laboratory Information Management System (LIMS), the data extraction and peak-identification software, data processing tools for QC and compound identification, and a collection of information interpretation and visualization tools for use by data analysts. The hardware and software foundations for these informatics components were the LAN backbone, and a database server running Oracle 10.2.0.1 Enterprise Edition."},

"MS_METABOLITE_DATA":{
"Units":"peak area",

"Data":[{"Metabolite":"(12 or 13)-methylmyristate (a15:0 or i15:0)","MICH-01419":"1.0081","MICH-01420":"0.6200","MICH-01421":"1.2463","MICH-01422":"0.3816","MICH-01423":"2.3662","MICH-01424":"1.2469","MICH-01425":"0.6884","MICH-01426":"0.2576","MICH-01427":"1.1514","MICH-01428":"0.5545","MICH-01429":"1.0019","MICH-01430":"1.0881","MICH-01431":"1.0461","MICH-01432":"1.0215","MICH-01433":"1.9402","MICH-01434":"1.1412","MICH-01435":"1.1713","MICH-01436":"0.6201","MICH-01437":"0.5445","MICH-01438":"0.8011","MICH-01439":"0.5412","MICH-01440":"0.4944","MICH-01441":"1.3022","MICH-01442":"1.3484","MICH-01443":"0.8400","MICH-01444":"0.8429","MICH-01445":"1.2588","MICH-01446":"1.0487","MICH-01447":"1.5621","MICH-01448":"1.6993","MICH-01449":"1.0402","MICH-01450":"0.9133","MICH-01451":"0.9690","MICH-01452":"0.5629","MICH-01453":"0.7815","MICH-01454":"1.1713","MICH-01455":"0.5235","MICH-01456":"0.8976","MICH-01457":"1.2710","MICH-01458":"1.6134","MICH-01459":"0.7299","MICH-01460":"0.6071","MICH-01461":"1.0749","MICH-01462":"0.9631","MICH-01463":"0.9312","MICH-01464":"0.9533","MICH-01465":"1.7222","MICH-01466":"0.7021","MICH-01467":"1.5460","MICH-01468":"0.6523","MICH-01469":"0.8054","MICH-01470":"1.6240","MICH-01471":"0.6574","MICH-01472":"0.9014","MICH-01473":"0.4519","MICH-01474":"1.0979","MICH-01475":"1.2957","MICH-01476":"1.2735","MICH-01477":"0.5803","MICH-01478":"2.4054","MICH-01479":"0.9635","MICH-01480":"0.7463","MICH-01481":"1.2640","MICH-01482":"1.1605","MICH-01483":"0.4697","MICH-01484":"0.8305","MICH-01485":"0.9964","MICH-01486":"0.5763","MICH-01487":"0.8392","MICH-01488":"0.8688","MICH-01489":"0.7456","MICH-01490":"2.6593","MICH-01491":"2.1402","MICH-01492":"0.8119","MICH-01493":"1.3559","MICH-01494":"1.0697","MICH-01495":"1.0332","MICH-01496":"1.2491","MICH-01497":"1.5458","MICH-01498":"0.7621","MICH-01499":"0.8186","MICH-01500":"1.7712","MICH-01501":"0.2576","MICH-01502":"1.3071","MICH-01503":"0.7597","MICH-01504":"0.2576","MICH-01505":"1.0020","MICH-01506":"0.8399","MICH-01507":"2.9332","MICH-01508":"1.7346","MICH-01509":"1.2557","MICH-01510":"1.1281","MICH-01511":"0.9981","MICH-01512":"0.7318","MICH-01513":"0.8153","MICH-01514":"1.5973","MICH-01515":"2.0693","MICH-01516":"0.9284","MICH-01517":"0.7026","MICH-01518":"0.8351","MICH-01519":"1.1646","MICH-01520":"0.6485","MICH-01521":"1.0575","MICH-01522":"1.2577","MICH-01523":"1.3420","MICH-01524":"0.5607"},{"Metabolite":"(14 or 15)-methylpalmitate (a17:0 or 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